Scientific Publications

Scientific Publications

(Click on the title for a link to paper)

Allen, M.L., H.U. Wittmer, E.P. Alexander, and C. Wilmers. In Press. Ontogeny of communication behaviors in a solitary felid. Behaviour.

Avrin, A.C., C. Pekins, J. Sperry, P. Wolff, and M.L. Allen. In Press. Efficacy of attractants for detecting eastern spotted skunks: an experimental approach. Wildlife Biology.

Baek, S., T. Iwasaki, K. Yamazaki, T. Naganuma, A. Inagaki, K. Tochigi, M.L. Allen, C. Kozakai, and S. Koike. In Press. Factors affecting predenning activity in Asian black bears. Mammal Study.

Bauder, J., N.M. Roberts, D. Ruid, B. Kohn, and M.L. Allen. In Press. Effects of translocation on survival of nuisance bears. Animal Conservation.

Bauder, J.M., M.L. Allen, T.J. Benson, C. Miller, and K. Stodola. In Press. An approach for using multiple indices for monitoring long-term trends of mesopredators at broad spatial scales. Biodiversity and Conservation.

Sebastián-González, E., Z. Morales-Reyes, F. Botella, L. Naves-Alegre, J.M. Pérez-García, P. Mateo-Tomás, P.P. Olea, M. Moleón, J.M. Barbosa, F. Hiraldo, E. Arrondo, J.A. Donázar, A. Cortés-Avizanda, N. Selva, S.A. Lambertucci, A. Bhattacharjee, A.L. Brewer, E. Abernethy, K. Turner, J.C. Beasley, T.L. DeVault, H.C. Gerke, O.E. Rhodes Jr, A. Ordiz, C. Wikenros, B. Zimmermann, P. Wabakken, C.C. Wilmers, J.A. Smith, C.J. Kendall, D. Ogada, E. Frehner, M.L. Allen, H.U. Wittmer, J.R.A. Butler, J.T. du Toit, A. Margalida, P. Oliva-Vidal, D. Wilson, K. Jerina, M. Krofel, R. Kostecke, R. Inger, E. Per, Y. Ayhan, M. Sanci, N. Ozturk, A. Inagaki, S. Koike, A. Samson, P.L. Perrig, E. Spencer, T.M. Newsome, M. Heurich, J.D. Anadon, E.R. Buechley, C. Gutierrez-Canovas, L.M. Elbroch, and J.A. Sánchez-Zapata. In Press. Functional traits driving species role in the structure of terrestrial vertebrate scavenger networks. Ecology.

78. Allen, M.L., L.M. Elbroch, and H.U. Wittmer. 2021. Can’t bear the competition: energetic losses from kleptoparasitism by a dominant scavenger may alter foraging behaviors of an apex predator. Basic and Applied Ecology 51: 1–10.

77. Allen, M.L., M. Sibarini and M. Krofel. 2021. Predicting preferred prey of Sumatran tigers (Panthera tigris sumatrae) via spatiotemporal overlap. Oryx 55: 197–203.

76. Allen, M.L., H.U. Wittmer, A. Ingaki, K. Yamazaki, and S. Koike. 2021. Food caching by bears: a literature review and new observations for Asiatic and American black bears. Ursus 32: e10.

75. Allen, M.L., L.M. Elbroch, and H.U. Wittmer. 2021. Scavenging by fishers in relation to season and other scavengers. Ecological Research 36: 353–359.

74. Allen, M.L., A.C. Avrin, M.J. Farmer, L.S. Whipple, E.P. Alexander, A.M. Cervantes, and J.M. Bauder. 2021. Limitations of current knowledge about the ecology of grey foxes hamper conservation efforts. The Journal of Threatened Taxa 13: 19079-19092.

73. Avrin, A.C., C. Pekins, J. Sperry, and M.L. Allen. 2021. Evaluating the efficacy and decay of lures for improving detections of carnivores with camera traps. Ecosphere 12: e03710.

72. Avrin, A.C., C. Pekins, and M.L. Allen. 2021. Eastern spotted skunk Spilogale putorius Linnaeus, 1758 (Mammalia: Carnivora: Mephitidae) mating behavior documented via camera trap. Journal of Threatened Taxa 13: 18660–18662.

71. Bauder, J.M., A.M. Cervantes, A.C. Avrin, L.S. Whipple, M.J. Farmer, C. Miller, T.J. Benson, K. Stodola, and M.L. Allen. 2021. Mismatched spatial scales limit the utility of citizen science data for estimating wildlife-habitat relationships. Ecological Research 36:87–96.

70. Cove, M.V., et al. 2021. SNAPSHOT USA 2019: the first coordinated national camera trap survey of the United States. Ecology 102: e03353.

69. Drake, D., S. Dubay, and M.L. Allen. 2021. Evaluating human-coyote encounters in an urban landscape using citizen science. Journal of Urban Ecology 7: juaa032.

68. Farmer, M.J. M.L. Allen, E.R. Olson, J. Van Stappen, and T.R. Van Deelen. 2021. Agonistic interactions and island biogeography as drivers of carnivore spatiotemporal activity at multiple scales in a temperate archipelago. Canadian Journal of Zoology 99: 309–317.

67. Jackson, K., C.C. Wilmers, H.U. Wittmer, and M.L. Allen. 2021. First documentation of scent marking behaviors in striped skunks (Mephitis mephitis). Mammal Research 66: 399–404.

66. Krofel, M., D. Juznic, and M.L. Allen. 2021. Scavenging and carcass caching behaviour by European wildcat, Felis silvestris. Ecological Research 36: 556–561.

65. Suraci, J.P., K.M. Gaynor, M.L. Allen, P. Alexander, J.S. Brashares, S. Cendejas-Zarelli, K. Crooks, L.M. Elbroch, T. Forester, A.M. Green, J. Haight, N. Harris, M. Hebblewhite, F. Isbell, B. Johnston, R. Kays, P.E. Lendrum, J. Lewis, A. McInturff, W. McShea, T.W. Murphy, M. Palmer, A. Parsons, M.A. Parsons, Mary E. Pendergast, C. Pekins, L. Prugh, K.A. Sager-Fradkin, S. Schuttler, C.H. Sekercioglu, B. Shepherd, L. Whipple, J. Whittington, G. Wittemyer, and C.C. Wilmers. 2021. Disturbance type and species life history predict mammal responses to humans. Global Change Biology 27: 3718–3731.

64. Wallace, C.F., J. M. Golla, and M.L. Allen. 2021. Direct observations of a wolverine scavenging at an active gray wolf kill. Northwestern Naturalist 102: 157–160.

63. Allen, M.L., S. Wang, L. Olson, Q. Li, and M. Krofel. 2020. Counting cats for conservation: seasonal estimates of leopard density and drivers of distribution in the Serengeti. Biodiversity and Conservation 29: 3591–3608.

62. Allen, M.L., N.M. Roberts, and J.M. Bauder. 2020. Relationships of catch-per-unit-effort metrics with abundance vary depending on sampling method and population trajectory. PLoS One 15: e0233444.

61. Allen, M.L., M. Sibarini, L. Utoyo, and M. Krofel. 2020. Terrestrial mammal community richness and temporal overlap between tigers and other carnivores in Bukit Barisan Selatan National Park, Sumatra. Animal Biodiversity and Conservation 43.1: 97–107.

60. Allen, M.L., A. Inagaki, and M.W. Ward. 2020. Cannibalism in raptors: a review. Journal of Raptor Research 54: 424–430.

59. Bauder, J., N.M. Roberts, D. Ruid, B. Kohn, and M.L. Allen. 2020. Nuisance black bear translocations across four decades indicates increased effectiveness by translocation distance and landscape context. Wildlife Research 47: 426–435.

58. Bauder, J.M., K. Stodola, T.J. Benson, C. Miller, and M.L. Allen. 2020. Raccoon pelt price and trapper harvest relationships are temporally inconsistent. Journal of Wildlife Management 1601–1610.

57. Bauder, J.M., M.L. Allen, T.J. Benson, C. Miller, A. Ahlers, and K. Stodola. 2020. Identifying and controlling for variation in canid trapper harvest data. Journal of Wildlife Management 84: 1234–1245.

56. Inagaki, A., M.L. Allen, T, Maruyama, K. Yamazaki, K. Tochigi, T. Naganuma, and K. Shinsuke. 2020. Vertebrate scavenger guild composition and utilization of carrion in an East Asian temperate forest. Ecology and Evolution DOI: 10.1002/ece3.5976.

55. Nickel, B. A., J. Suraci, M.L. Allen, and C.C. Wilmers. 2020. Human presence and human footprint have non-equivalent effects on wildlife spatiotemporal habitat use. Biological Conservation 241: 108383.

54. Sebastián-González, et al. 2020. Network structure of vertebrate scavenger assemblages is driven by ecosystem productivity and human impact at a global scale. Ecography: 43: 1143–1155.

53. Yovovich, V., M.L. Allen, L. Macauley and C.C. Wilmers. 2020. Using spatial characteristics of apex carnivore reproductive behaviors to inform conservation planning. Biodiversity and Conservation 29: 2589-2603.

52. Olson, L.O., T.R. Van Deelen, J.D.J. Clare, and M.L. Allen. 2020. Proximity to established populations explains moose (Alces alces) occupancy in northern Wisconsin. American Midland Naturalist 183: 24–37.

51. Allen, M.L., R.E. Harris, L. Olson, E. Olson, J. Van Stappen, T.R. Van Deelen. 2019. Resource limitations and competitive interactions affect carnivore community composition at different ecological scales in a temperate island system. Mammalia 83: 552–561.

50. Allen, M.L., M.P. Ward, D. Južnič, and M. Krofel. 2019. Scavenging by owls: a global review and new observations from Europe and North America. Journal of Raptor Research 53: 410–418.

49. Allen, M.L., N.M. Roberts, M.J. Morales, and T.R. Van Deelen. 2019. Decreasing available bobcat tags appear to have increased success, interest, and participation among hunters. Human Dimensions of Wildlife 24: 349-362.

48. Allen, M.L., C. Norcross, and P. Perrig. 2019.Puma (Puma concolor) predation on novel pinniped prey. Northwestern Naturalist 100: 209–211.

47. Allen, M.L., M.C. Sibarani, and L. Utoyo. 2019. First record of a wild hypopigmented oriental small-clawed otter (Amblonyx cinereus). Ecotropica  21: 201904.

46. Allen, M.L., and S.M. Crimmins. 2019. Sea otters from the northern and southern populations may find each other in Humboldt County, California. IUCN Otter Specialist Group Bulletin 36: 88–92.

45. Allen, M.L., L. Hocevar, and M. Krofel. 2019. Resource Holding Potential. In: Encyclopedia of Animal Cognition and Behavior, eds: J. Vonk and T.K. Shackleford. Springer, New York.

44. Mueller, M.M., D. Drake, and M.L. Allen. 2019. Using citizen science to inform urban canid management in Madison, Wisconsin. Landscape and Urban Planning 189: 362-371.

43. Cristescu, B. S. Bose, L.M. Elbroch, M.L. Allen, and H.U. Wittmer. 2019. Habitat selection when killing primary versus alternative prey species supports prey specialisation in an apex predator. Journal of Zoology 309:259-268.

42. Farmer, M.J. and M.L. Allen. 2019. Persistence in the face of change: effects of human recreation on coyote (Canis latrans) habitat use in an altered ecosystem. Urban Naturalist 29: 1–14.

41. Olson, L.O. and M.L. Allen. 2019. A leucistic fisher (Pekania pennanti) and the prevalence of leucism in wild carnivores. American Midland Naturalist 181: 133–138.

40. Sebastián-González, E., et al. 2019. Scavenging in the Anthropocene: human impact drives vertebrate scavenger species richness on a global scale. Global Change Biology 25: 30053017.

39. Allen, M.L., A.S. Norton, G. Stauffer, N.M. Roberts, Y. Luo, Q. Li, D. MacFarland, and T.R. Van Deelen. 2018. A Bayesian state-space model using age-at-harvest data for estimating the population of black bears (Ursus americanus) in Wisconsin. Scientific Reports 8: 12440.

38. Allen, M.L., N.M. Roberts, and T.R. Van Deelen. 2018. Hunter selection for larger and older male bobcats affects annual harvest demography. Royal Society Open Science 5: 180668.

37. Allen, M.L., B. Peterson, and M. Krofel. 2018. No respect for apex carnivores: Distribution and activity patterns of honey badgers in the Serengeti. Mammalian Biology 89: 90–94.

36. Allen, M.L., M.J. Morales, E.R. Olson, J. Van Stappen, and T.R. Van Deelen. 2018. Is there anybody out there? Occupancy of the carnivore guild in a remote temperate archipelago. Community Ecology 19: 272–280.

35. Allen, M.L. N.M. Roberts, and T.R. Van Deelen. 2018. Age-at-harvest models as monitoring and harvest management tools for Wisconsin carnivores. Final Report to the Wisconsin Department of Natural Resources.

34. Allen, M.L., B. Wojcik, B.E. Evans, E.E. Iehl, R.E. Harris, M.E. Wheeler, B.E. Peterson, R.L. Dohm, M.A. Mueller, L. Olson, B. Ederer, M. Stewart, S. Crimmins, K. Pemble, E. Olson, J. Van Stappen, T.R. Van Deelen. 2018. Detection of endangered American martens (Martes americana) in Apostle Islands National Lakeshore, Wisconsin. American Midland Naturalist 179: 294–298.

33. Mueller, M.M., D. Drake, and M.L. Allen. 2018. Coexistence of coyotes (Canis latrans) and red foxes (Vulpes Vulpes) in an urban landscape. PLoS One 13: e0190971.

32. Allen, M.L., L. Hocevar, and M. Krofel. 2017. Where to leave a message? The selection and adaptive significance of scent-marking sites for Eurasian lynx. Behavioral Ecology and Sociobiology 71: 136.

31. Allen, M.L., M.S. Gunther and C.C. Wilmers. 2017. The scent of your enemy is my friend? The acquisition of large carnivore scent by a smaller carnivore. Journal of Ethology 35: 13–19.

30. Allen, M.L., B. Kohn, N.M. Roberts, S.M. Crimmins, and T.R. Van Deelen. 2017. Benefits and drawbacks of determining litter success for black bears (Ursus americanus) from cementum annuli techniques. Canadian Journal of Zoology 95: 991-995.

29. Allen, M.L., M.J. Morales, M.E. Wheeler, J.D. Clare, M.A. Mueller, L.O. Olson, K. Pemble, E.R. Olson, J. Van Stappen, and T.R. Van Deelen. 2017. Survey techniques for determining distribution, abundance, and occupancy the carnivore guild in the Apostle Islands National Lakeshore (2014-2017). Final Report to the National Park Service.

28. Krofel, M., L. Hocevar, and M.L. Allen. 2017. Does human infrastructure shape scent marking in a solitary felid? Mammalian Biology 87: 36-39.

27. Chiang, P.-J., and M.L. Allen. 2017. Literature review of our current knowledge of clouded leopards. International Journal of Avian and Wildlife Biology 2: 00032.

26. Allen, M.L., C.C. Wilmers, L.M. Elbroch, J.M. Golla, and H.U. Wittmer. 2016. The importance of motivation, weapons and foul odors in driving encounter competition in carnivores. Ecology 97: 1905–1912.

25. Allen, M.L., H.U. Wittmer, E. Setiawan, S. Jaffe, and A.J. Marshall. 2016. Scent marking in Sunda clouded leopards (Neofelis diardi): novel observations close a key gap in understanding felid communication behaviours. Scientific Reports 6: 35433.

24. Allen, M.L., V. Yovovich, and C.C. Wilmers. 2016. Evaluating the responses of a territorial solitary carnivore to potential mates and competitors. Scientific Reports 6: 27257.

23. Allen, M.L., Y. Wang, and C.C. Wilmers. 2016. Exploring the adaptive significance of five types of puma (Puma concolor) vocalizations. Canadian Field-Naturalist.

22. Allen, M. L., B. E. Evans, M. E. Wheeler, M. A. Mueller, K. Pemble, E. R. Olson, J. Van Stappen, and T. R. Van Deelen. 2016. Survey techniques, detection probabilities, and the relative abundance of the carnivore guild on the Apostle Islands (2014-2016). Final Report to the National Park Service.

21. Allen, M.L., L.M. Elbroch, C.C. Wilmers, and H.U. Wittmer. 2015. The comparative effects of large carnivores on the acquisition of carrion by scavengers. American Naturalist 185: 822–833.

20. Allen, M.L., H.U. Wittmer, P. Houghtaling, J. Smith, L.M. Elbroch, and C.C. Wilmers. 2015. The role of scent marking in mate selection by female pumas (Puma concolor). PLoS One 10: e0139087.

19. Allen, M.L., C.F. Wallace, and C.C. Wilmers. 2015. Patterns in bobcat (Lynx rufus) scent marking and communication behaviors. Journal of Ethology 33: 9–14.

18. Allen, M.L., L.M. Elbroch, D.S. Casady, and H.U. Wittmer. 2015. The feeding and spatial ecology of mountain lions (Puma concolor) in Mendocino National Forest, California. California Fish and Game Journal 101: 51–65.

17. Allen, M.L., B.E. Evans, and M.S. Gunther. 2015. A potential range expansion of the coastal fisher (Pekania pennanti) population in California. California Fish and Game 101: 280-285.

16. Wang, Y., M.L. Allen, and C.C. Wilmers. 2015. Mesopredator spatial and temporal responses to large predators and human development in the Santa Cruz Mountains of California. Biological Conservation 190: 23–33.

15. Elbroch, L.M., P. Lendrum, M.L. Allen, and H.U. Wittmer. 2015. Nowhere to hide: pumas, black bears, and competition refuges. Behavioral Ecology 26: 247–254.

14. Taylor, A.P., M.L. Allen, and M. Gunther. 2015. Marking behavior on rub trees by black bears during breeding season. Behaviour 152: 1097-1111.

13. Allen, M.L. 2014. The Ecology and Behaviour of Pumas (Puma concolor) in Northern California. Ph.D. Dissertation, Victoria University, Wellington.

12. Allen, M.L., L.M. Elbroch, C.C. Wilmers, and H.U. Wittmer. 2014. Trophic facilitation or limitation?Comparative effects of pumas and black bears on the scavenger community. PLoS One 9: e102257.

11. Allen, M.L., H.U. Wittmer, and C.C. Wilmers. 2014. Puma scrape and communication behaviors: understanding functional use and variation by sex and age. Behaviour 151: 819–840.

10. Allen, M.L., L.M. Elbroch, D.S. Casady, and H.U. Wittmer. 2014. Seasonal variation in the feeding ecology of pumas (Puma concolor) in northern California. Canadian Journal of Zoology 92: 397–403.

9. Elbroch, L.M., M.L. Allen, B.H. Lowery, and H.U. Wittmer. 2014. The difference between killing and eating: ecological shortcomings of puma energetic models. Ecosphere 5: article 53.

8. Wittmer, H.U., T.F. Forrester, M.L. Allen, L. Marescot, and D.S. Casady. 2014. Black-tailed deer population assessment in the Mendocino National Forest, California. Final Project Report, California Department of Fish and Wildlife.

7. Allen, M.L., L.M. Elbroch, and H.U. Wittmer. 2013. Encounter competition between a Cougar, Puma concolor, and a Western Spotted Skunk, Spilogale gracilis. Canadian Field-Naturalist 127: 64–66.

6. Allen, M.L., and A.P. Taylor. 2013. First record of scavenging in a western screech owl (Megascops kennicottii). Wilson Journal of Ornithology 125: 417–419.

5. Wilmers, C.C., Y. Wang, B. Nickel, P. Houghtaling, Y. Shakeri, M.L. Allen, J. Kermish-Wells, V. Yovovich, and T. Williams. 2013. Scale dependent behavioral responses to human development by a large predator, the puma. PLoS One 8: e60590.

4. Casady, D.S., and M.L. Allen. 2013. Handling adjustments to reduce chemical capture–related mortality in black-tailed deer. California Fish and Game 99: 104–109.

3. Elbroch, L.M., and M.L. Allen. 2013. Prey indices and behaviors at a gray fox den in San Mateo County, California. Western North American Naturalist 73: 240–243.

2. Allen, M.L. 2013. Scavenging of a striped skunk (Mephitis mephitis) by a ringtail (Bassariscus astutus) in northern California. Northwestern Naturalist 94: 79–81.

1. Allen, M.L., and D.S. Casady. 2012. Recent observations of porcupines in El Dorado County, California. California Fish and Game 98: 175–177.